"Frequently, the internal domain name differs from the external domain name. For example, your internal domain name may be domain.local, although your external domain name is domain.com. The domain name system (DNS) MX resource record for your organization references domain.com. Domain.com is the SMTP namespace that you assign to users when you create an e-mail address policy. You must create an accepted domain to match your external domain name."
1. Open Exchange Management Console -> expand Organization Configuration -> click on Hub Transport.
2. Go to Accepted Domains tab -> New Accepted Domain.
3. Go to E-mail Address Policies -> New E-mail Address Policy.
Useful links:
http://technet.microsoft.com/en-us/library/bb124423%28EXCHG.80%29.aspx
http://blogs.technet.com/b/exchange/archive/2006/11/16/3397300.aspx
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